IS3

PHOTO SYNTHBSLS IN AN
tant
peen plantt 3
puotoaynthass Food 2 O2 CPoduc"
einngy all
a C og depeud! upon uniglht ton eringy

anly xperuMents houad

Catgop hyu Lught co 0a fo HTSN (Photeana)
2 kauh tns n
ganpants ncs u g b
tpaxd pan
3Erposed
al teskd shanehte
C O s 4: fpbetosynthess

eepenmen
lanly

Teoph ueetkey C930) esBential o k ain an 9-ouh,

dis counnod O2
MOUL kocated whon plaLod un j a uth ardle
ouut Las aluine when mint plant aluo kept 2

candls alas kept on kwning.

Plans ushat braatuig anma 2

condlos ILmBIL,

the milaon aetup ente u

Jan 9ngenhous blacead
sUntoht
Subsus
h unlught ; Amal
9LAen pbans taile n dank t y did not
Thsa lbbes oXvOE
'untug he s euential 2 ouly ten parta con suLaa se

n plon
Julus Ven soehs ( S ) qucas poodut"
chtoneb tn chlorsfplosts.
n qpaan bans
twe qluege STARCH

TW engelman Pouism 4peub gut3 4beutral cempo

pAad in 4bens Len.
WLns Lluuninated ladephera
aenobic basenia rean alga)
O, euoluhen.
datetted
Bauleua mainy luu Sud gle
aeuunutated

Achen peetrumus dexibed 4 Aesembla

chlgnophyU a ab
ablopbom
KEY FEATURES kniowN wmid 19" Conhny
b
plan us linht to make Conbob
gt
CO H
Apn elius van NOl puuplu ga2n bautenNa S home
PHSTSIS aht dapendaut on in which
Hrom ondis able cemb oduca D to CHO

HA t C0 ught
A tCH,Otho

G gheen plouet ho donan ems

erm ud not Co. (po uad ky nadioiuoh

.comeet e- tmuth pmas)
4C0 t l& H,o ght C,+6ho t60,
LgueoLey
wWhne does photoiyntheis tale place
=

n n n leauAs u aso n othen

graon panrs. ftm)
LewLs mopyl celll lag no chlonsplasbs
algn hwmew alorg uall mezophyl ge
openumuht)
n chlonoplazs
LyHT RNS membans ut. tnop enony
(photo chanias an) ynthosde TP NADPH

pAR BNs Lt uelependat Y depeudantJon

pHaduek
n Stremg
eyymih men 4L oUms
 Puguuen
uwotund n Reuotpynthesi
colon n aus du to
ehleropbytl a
Cbugut1 6ua qruan)
chiorOphye b
-

Cyellou gan)
Xauthaphyts Cqulou
Canatenoidg
llow -00nau )
men
ans aku
abiluty to abcoub ualut at peufiu usawlangeas
Laus laL b &rd

Camenod

Asop ae
P'gmen

phoos

A BSORPrION CPecTOM ACtDN ePezTROu

le ahiop") (OLnt o absorb d )

pgment auten

chlooplyela tua (hie pig nan to ph0toryn 2

mar" abionp" tole plau. in ud ragi ons.

at otsh A but lassen

(&ome phoos yn. also tae plaLe
chl'b,Xauthophyls
CaenA acersey PLgnunk.
aboub ugut
thansA to chA'a
prolee che'a ro
photoOxtdan
IGHT KXN
C PAOkoeuenucal phae)
Undudos i g h t aboypt
alitt ing
0 oleose
3 olaase o ATP 3 AADPH

u
LHC tntE(igh hoitung complees) Ps I PS I
(6ep nm) 700 nin)
LHGLHG hag wany Pgmnts bound to (wuin.
Rxn canhe chs. 'a
Ansennae othe pgwants chs b, canskn OUnthe

antehna
Ccha'a)

Antennat herp tn me
ctient photoa H ablonbu
Light didiy As.
Rn cenren
s topD
P6&o
Eleton
Tuuor
P abeenb
n A ad es eued
junp
hLen onbtr Passed emansportyaem cesitig-
cy6ockroms
Cotouonuand meuwnt es n teams o{ edox
potntial Sak
NADP
eaccept ATP a ceupo

LHE) Canhit

PSTL

Z-Scheme
gut sn
PST miu) aUep or dlounn) 6TS PS
CdousnillNADr oms ADPH +HT (phiu) accu

Covw rom, H Coly exetad

at PSL )(Unnur áde
Ho 4H+0 +4e turaiecid muwb
hsphPlaten ATP ntihssis (nnitohondaia 2 chteroptl
PaGLphaLoyahon ATPdy nth. n trte ignt
(re ADr ae ) APP4 ATP ]

NOn ucuc plthopkephovylation Uwn both PsA PSI wmk

2 scherme)
Cydie PST funt "al
in stoma mellae Cas theu
tok ps a n them NADPKaduuhau
C3yme)
0ndy ATP iythaszed
these alss rcn
aA only4 auailasle. n
tuytated membyant.

e acup08

LHC

PsI

CHE MDEMOTIC HYPOTHESL

ATP n h inAed to del pofon quaoient avDu merb. o
bhuykako.
Ht acouwmulated touuDvds nlds a tylakoid ( twma)

Lin rospi", i n tespau et mitvehon. embavn).

Popton gradent deuulope duu to i
H 0 plttvg Htaceum h luman
e mout trough PS .
Ht acumulate tnsde bcoz e
thnoun tounands e acceptor 2 hunes probn omwuld
Om tToma
3 9 NADP Suduuhase enz4me r e g . H on redue , H' J s un
srorma.
Cin crona
 2 u lumen Tde
H in shwrma
se 3 Measons
Mhi ca t u a hsoton quodient,

ths gradeut n. U
LE atoLun o srrorna
(Luynen -
ATP ynthase CE)
enme

cfo
embedded n
c (0 es touuands

thytax. mumb bemaa

hen H MLU ) umen

confiq chang n Cf n e3ed Har

manu ATP.

STeoM ADP AT
NADPNADPH

sadiant
Cyroch
oma
B6
Piastoqin

PS I Ptaztoay PST
tg-
adunt

WMON H
tigh HT)

Chumoa/mous , ATP Hnthase has a

membvane
ehainel that allos
dilhus
H Punp
Potens bauk to roa
proron 9Hadent Hu llease enwe
ATP mtase ATP.

Lsed n
bYug co2

Pasus ght At, NAD'H, 0

 whene N_ATP& NmDPH d
ut 3 Produeu
dituses ou og chlonoplatt
ATe& NADPH
Sugan lutot ynth,
Then e0mes he
oynt hetic phase,
labu u
phasu ontinuni {on wme timw hun
Stops,
aLaitable Syn. stasks again.
Callad DaK Un Caeas not it taks
Mian
plau ih dank

Thu takas plae

CCHD) t o, J

Meuin Caluh Cused algal phoro qn. studu

He shudiud th 4 shabu pModuutr n canben ffasi on
kends shabl pralutks uwn ound.

3PUA Cy eatuin) OAA Chy oehoru scientist)

3-Phaphogeyuric auid ovale autie aiid
3 cabon comp (4-c-omfowd ) t

Ca pathuLa C patlhuay

Pumay accepo oCO2 2RuGP s-c)

CRlbulou olsphosphaiu

tanlien, they expected acepo to be 2-c comp' PqA a

CALVIN CYLA
KuBP sageunnted.
Calun yeu a u phetoeyntheit plant
ie bothn
L cauln eyely
O Canboxylat2 Replus Kegennahon
 NAD'/Npof 4H NAOv
(oyded (eduued
pwm)

conboxylase Cezyma
atmutty othan than canborylase
Aue,alto Aas @Xygenathen
cwnAtnt to call t koB Co
9t moa

Cauin cyele
(Aurnosphent)

RuBP
C0,+ H2o
RuBo
eaboxylation

ADPF Keqeneatioi ATP

3PGA
ATP NADPH
Redutthen NAPP

Tie
Phetphate

n one cycla, co moleule bixa)

3PG med
ATP 2 NADPH Useol Cin redua")

4ATP wed ( rgenara")

C 2 NADDH 3 3ATP req n on
y cRe
TO om 4 HO) moleulL
6 tuwns
Hde a
ie.CCc02. mouuls
9n Out

6 CO2 qdeucost
18 ATP 18 ADP
12 NADPH 12NApP

gn (3 ?hokorepua tas -kiva lce

C4 Photorespuat
 hauo.
16
ha RUBisCO &ymo, has rtlcaLe
Kundle

M&CD, Rack
tf cas
PATHWAy

d opual
Lin" stahle ugiou
P l a u

-
Cy pakuwau
produet 0AR
thuein man uptuyutheeie pcthuta
Cy plen peauial
n tolenoute Weaanatemy Kron anatoj
high emp hgh
Jauk pbnowdapinoan unue uveni
guuaten
puodutuudy
Knar ahat
om CA hau Lwndlu dheath calls ulth
(unuath)
chlonoplas anouLnd s . haue iek als & ne
fnterceulan Spaces

Mauze Soglum 00go- Meopklt

6 S cells

Capathuuny atuo catted Hatch &dlack Pathuuay

nimayy aepro co PeP CPhepheenDl pyuvat
Cin muophyu)

Atm CO2

HCO+ PEP PEP

Par
caNoxylas
FIXRTION
NALOphy REMENERATIONJ
OAA C4¢)
uwate

Mali aid cq-

aspantic acid

Natic aed

&upd
DECARO0XVLATION

PyBLLVate

Co, CLe) (ac

calin yode
 catun yetu n mesophyy
bundl healh

PHOTOREPPIRATIO N
gut exn

ATe
NADPH omed

LRuslo (rou abunuty o O)

high
Co+ RuBCo O2 t Rugisp

CALVIN CHLE HOTORESPPATLON

Conbohydnatu V C Canbolyd. X

Pugisco
KuBP + 0 2x(3P4A
Ku'st0 m0&t abllhdant engyme in tho uSnid

Cajual hen
ueate inity to CD_
n 2 bunds to RuBP
Co ix su.
RuBP+0, PGA+ PGO
3C) (2
Cphaphagywate) CPLSphoglycolata)
No AT 2 suga ntsU ake pla te N4DPH alo
Rathen uLes A TP 2 alenses Co
 163

atFato 41eetng Photalyuthesis

Rata plotoyn >
any unp otermina
to uyuld produts,
odapends on auos botth exeuna t veunal)

untuglt, temp (gonakie redispoel Hon 2

Co cenu, H gkauLth plant)

Blackman's lau imdt'hg faeors 4 a chamisa uoco

eeted hy 1 ormonfacto thon a e s detminad by
acto cuch nearest 6o it mininal ualur,

A GH7
Thee s d n eht quali, ebt itenuit Light pesus
At low ineniu unean w xate.

hg ntensuk gkaduatly. no
Encrase

igtr satunath occuns at 1o

uel unltigbe, Lughe unt¬nsu
ht aNey a unitig facton

Incxaase aintensit causes buakdauun etlonphyl

e in a t e ,

CO comaentration

Majon umrin9 faton,

6 atmn Co conc O 03 0-04 , )
but reiuts ans damagup
Beyond0 05 cokra raks

C C4 s0spemd denanty.
Czat
C
LOu CO,
se
(260pt satuna")
C450 ul sat")
Limiting
Cused nhausu
pps fomaro
 TEMPORATU PE
ENZMATIC temb couhmLled
Dank Jen

C Cy
low tmp

Hugh+emp tse
Habitut also datenmine ehtimum emp
Tovopieat plants haua Alhey tomp epimun
than emporateclimats.

WATER
dres cnuses stvnuata 4o dlose phshe,
cO2 aulabltt
D Itre alus_makes Jeaues u l t , seye Hhe
Ayate anea
leaues j Jug metabolie actuuity.